The high level connection (speaker level) at your sub is in essence a parallel connection. One (to your speakers) incorporates a high pass filter, like a speaker crossover.
The other ... the signal that the sub uses, will pass through a transformer that changes it to a line level signal, which is then used by the sub just as if it were line level (RCA) input from a preamp. All the signal processing the sub performs is the same whether it's a line level input or a speaker level input.